- Peter Cullen voices Autobot leader Optimus Prime. He retains his alternate mode of a blue Peterbilt truck with red flame decals. Cullen recorded a voiceover for the opening scene in August 2008, but began the majority of voice work in November. A live action cameo has been written for Cullen.
- Mark Ryan voices Bumblebee the Autobot who befriended Sam and disguises himself as his fifth-generation Chevrolet Camaro. Ryan also continues his role as stand-in for the robots on set.The filmmakers used the Super Sport version of the 2010 Camaro rather than the concept version. Ed Welburn, vice president of GM Global Design, said the redesigned car emphasized Bumblebee as becoming stronger after having his severed legs reattached in the first film: the new intakes and spoilers show he is a sturdier character.
- Ironhide, the Autobot weapons specialist who transforms into a GMC Topkick. Jess Harnell voiced him in the first film, but it is unknown whether he signed to reprise the role.
- Ratchet, the Autobot medic who transforms into a search and rescue Hummer H2. Robert Foxworth voiced him in the first film, but it is unknown whether he signed to reprise the role.
- Mudflap and Skids, respectively a red Chevrolet Trax and green Chevrolet Beat. Mudflap is very hyperactive, while Skids believes himself to be the smarter of the two and tries to come across as mature, but nevertheless appears unable to keep quiet. Bay chose the Beat and Trax for the Autobot twins because he thought those two small cars looked good together. Ed Welburn added when they had designed the cars before Bay selected them, the "character" they wanted to bring to the concepts was a sense that small cars could be as cool as large ones. According to leaked daily call sheets that Bay claimed he faked,They begin the film by combining into an ice cream van. Skids was mentioned by the other Autobots in the Transformers: The Reign of Starscream movie sequel comics.
- Sideswipe, a silver Chevrolet Corvette Stingray Concept.
- Jolt, a blue Chevrolet Volt. Jolt was a last-minute addition to the cast after General Motors, the film's car supplier, wanted to promote the Volt. The writers had already wanted to include the car in the script before the Writers Guild strike, so they had to work out a character that would fit well within the Autobot team afterwards and convince Bay to approve the addition. Welburn was pleased the Volt's character was christened Jolt by the filmmakers, as it reflected that car's design "character".
- Arcee, an Autobot with a feminine personality and appearance, who can separate into three pink, purple and light blue motorcycles with their own robot forms. Arcee was dropped from the first film as the writers felt there was not enough time to explain her "gender", but she and several other female Transformers appeared in the first film's toy line and tie-in comics. Orci stated the issue of gender could just be ignored in the film.
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